Brillouin Index Calculator

The Brillouin Index Calculator measures species diversity for complete census data (when every individual in a community is counted). Enter your species counts — the number of individuals for each species — and the tool computes the Brillouin Index (H), along with maximum possible diversity (H_max) and evenness (E). Unlike Shannon or Simpson indices, Brillouin is the correct choice when you have a full enumeration rather than a sample.

Enter the total count of individuals for this species.

Results

Brillouin Index (H)

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Maximum Diversity (H_max)

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Evenness (E = H / H_max)

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Total Individuals (N)

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Species Richness (S)

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Individual Counts per Species

Results Table

Frequently Asked Questions

What is the Brillouin Index?

The Brillouin Index (H) is a measure of species diversity developed by Louis Brillouin. Unlike the Shannon Index, it is specifically designed for situations where you have conducted a complete census of a community — meaning every individual has been counted — rather than working with a sample. It accounts for the exact number of individuals rather than estimated proportions.

What is the formula for the Brillouin Index?

The Brillouin Index is calculated as H = (1/N) × ln(N! / ∏n_i!), where N is the total number of individuals and n_i is the count for each species. Because factorials of large numbers are involved, the calculation uses the log-gamma function (Stirling's approximation or summing logarithms) for numerical stability.

When should I use the Brillouin Index instead of the Shannon Index?

Use the Brillouin Index when your data represents a complete census — every individual in the community has been counted and identified. Use the Shannon Index when you are working with a random sample from a larger, unknown population. Using the Shannon Index on census data can introduce a small bias that Brillouin corrects for.

What does a high Brillouin Index value indicate?

A higher H value indicates greater diversity — the community contains many species distributed relatively evenly across individuals. A value near H_max means the population is as diverse as it could be given the number of species and total individuals present.

What does evenness (E) mean in this context?

Evenness (E = H / H_max) measures how close the observed diversity is to the theoretical maximum. A value of 1.0 means all species are represented equally. Values closer to 0 indicate that one or a few species dominate the community, even if many species are present.

What is H_max in the Brillouin Index?

H_max is the maximum possible Brillouin diversity for a community with the same total number of individuals (N) and the same number of species (S). It is computed assuming individuals are distributed as evenly as possible across all species. It serves as a benchmark for interpreting how diverse the observed community is.

How many species can I enter in this calculator?

This calculator supports up to 8 species. Enter a count of 0 (or leave blank) for any species slot you do not need — those entries will be automatically excluded from the calculation. For studies with more species, the same formula applies and can be extended manually.

Does the Brillouin Index work for very small communities?

Yes, but the index becomes less informative when N (total individuals) is very small (e.g. fewer than 5). The factorial-based formula is mathematically exact for any N, but ecologically meaningful interpretation generally requires a reasonably sized community with at least several individuals per species.

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